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FOSSIL 2: GASTROPODA AND CEPHALOPODA

The next types of fossils are those of gastropoda and cephalopoda. When describing fossils of these two groups, additional terms are often used: trochospiral and planispiral.


Trochospiral is usually associated with gastropoda and refers to a three-dimensional coiling of the shell, meaning the shell has height and spirals from top to bottom. There are two types of coiling directions in trochospiral forms: dextral and sinistral. A dextral coil spirals in a clockwise direction, while a sinistral coil spirals counterclockwise.


To make it easier to understand, you can look at the position of the aperture (the opening where the gastropoda emerges). If the aperture is on the right side, the coiling is dextral; if it’s on the left side, the coiling is sinistral.



Sinistral (left) and Dextral (right)
 
Planispiral is typically found in cephalopoda and refers to a two-dimensional shell coiling (lacking height). In planispiral forms, there are three types of coiling patterns: involute, convolute, and evolute.


GASTROPODA

A. Definition

  • EtymologyGaster = stomach and Podos = foot
  • Terminology: Gastropoda are soft-bodied animals that move using a muscular “foot” located on their underside (belly).

B. Characteristics

  • Possess a chewing organ located in the head region, known as the mantle cavity.
  • Their shells are composed of calcium carbonate (CaCO₃).
  • The shell typically has a conical shape.

C. Taxonomic Level

  • Class: Gastropoda

D. Order Classification

a. Subclass Protogastropoda, consisting of the orders:

  • Cynostraca
  • Cochliostracea

b. Subclass Prosobranchia, consisting of the orders:

  • Archaeogastropoda
  • Mesogastropoda
  • Neogastropoda

c. Subclass Opisthobranchia, consisting of the orders:

  • Pleurocoela
  • Pteropoda

d. Subclass Pulmonata, consisting of the orders:

  • Basommothopora
  • Stylommothopora

E. Morphology



F. Index Fossil

  • Vicarya verneulli vicassa: Middle Miocene (N13) and found in the North Pacific region.


  • Turricula flammea atjehensis: Plio-Pleistocene, serves as evidence of Java-Sumatra tectonics during the Tertiary period.
  • Babylonia gracilis & Babylonia pangkaensis: Plio-Pleistocene.

Babylonia gracillis 
Babylonia pangkaensis

G. Oostingh’s Biozonation

In tropical regions, gastropods thrive exceptionally well, and certain gastropod species are characteristic of specific layers. This fact was used by Oostingh to establish the Neogene stratigraphy of Java Island. The biozones are as follows:

a. Rembang Stage (Early Miocene)

  • Index fossil: Turritella subulata

b. Preanger Stage (Middle Miocene)

  • Index fossil: Turritella angulata

c. Cirodeng Stage (Late Miocene)

  • Index fossil: Turritella angulata cramatensis

d. Cirebon Stage (Early Pliocene)

  • Index fossil: Turritella angulata acuticarinata

e. Sunda Stage (Late Pliocene)

  • Index fossil: Turritella angulata tjicumpalensis

f. Banten Stage (Early Pleistocene)

  • Index fossil: Turritella angulata bantamensis



CEPHALOPODA


A. Definition

  • Etymology: Cephale = head and Podos = foot
  • Terminology: Mollusks whose heads are surrounded by feet that have been modified into tentacles.

B. Characteristics

  • Generally have internal shells (like belemnites), but some have external shells (such as nautiloids and ammonoids).


Belemnites

Ammonoids

  • They release ink as a defense mechanism.

C. Taxonomic Level

  • Class: Cephalopoda

D. Morphology




E. Evolution of Suture

  • Sutures in cephalopoda can be used as an indicator of fossil age because their sutures evolved from simple forms to more complex patterns over time.

Goniatites

Cerratitics

Ammonites

  • Goniatites: indicator of the Triassic period.
  • Ceratitics: indicator of the Jurassic period.
  • Ammonites: indicator of the Cretaceous period.

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